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Molokaʻi is home to a wingless fly among many other endemic insects.

It used to be thought that Molokaʻi was first settled around AD 650 by indigenous peoples most likely from the Marquesas Islands.

However, a 2010 study using revised, high-precision radiocarbon dating based on more reliable samples has established that the period of eastern Polynesian colonization of the Marquesas Islands took place much later, in a shorter time frame of two waves: the "earliest in the Society Islands Later migrants likely came from Tahiti and other south Pacific islands.

Although Captain James Cook recorded sighting Molokaʻi in 1778, the first European sailor to visit the island was Captain George Dixon of the British Royal Navy in 1786.

The total 2010 census population of these was 7,345, Molokaʻi is split into two main geographical areas.

Below 4,000 feet (1,200 m), the vegetation is dominated by exotic flora, including strawberry guava (Psidium littorale), eucalyptus (Eucalyptus spp.), and cypress (Cupressus spp.).It lacks significant ground cover and virtually the entire section is covered in non-native kiawe (Prosopis pallida) trees.One of the few natural areas remaining almost intact are the coastal dunes of Moʻomomi, which are part of a Nature Conservancy preserve.The first significant European influence came in 1832 when a Protestant mission was established at Kaluaʻaha on the East End of the island by Reverend Harvey Hitchcock.The first farmer on Molokaʻi to grow, produce and mill sugar and coffee commercially was Rudolph Wilhelm Meyer, an immigrant from Germany who arrived in 1850.

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He built the first and only sugar mill on the island in 1878, which is now a museum.